Anhangueridae
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea.[1] They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested Tropeognathus.[2]
| Anhanguerids Temporal range: Early-Late Cretaceous,   | |
|---|---|
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| Anhanguera santanae skeleton | |
| Scientific classification  | |
| Kingdom: | Animalia | 
| Phylum: | Chordata | 
| Order: | †Pterosauria | 
| Suborder: | †Pterodactyloidea | 
| Clade: | †Anhangueria | 
| Family: | †Anhangueridae Campos & Kellner, 1985  | 
| Type species | |
| †Anhanguera blittersdorffi Campos & Kellner, 1985  | |
| Subgroups | |
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Relationships
    
There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set (Andres, 2021).[3]
| Anhangueridae | 
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The cladogram below follows Pêgas et al. (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while Ornithocheirus itself was recovered as a basal member of Ornithocheirae.[4]
| Anhangueria | 
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References
    
- Campos D. A.; Kellner Alexander W. A. (1985). "Panorama of the flying reptiles study in Brazil and South America". Anais da Academia Brasileira de Ciências. 57: 453–466.
 - Rodrigues, Taissa; Kellner, Alexander W. A. (2013). "Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England". ZooKeys (308): 1–112. doi:10.3897/zookeys.308.5559. PMC 3689139. PMID 23794925.
 - Andres, B. (2021) Phylogenetic systematics of Quetzalcoatlus Lawson 1975 (Pterodactyloidea: Azhdarchoidea). Journal of Vertebrate Paleontology, 41:sup1, 203-217. DOI: 10.1080/02724634.2020.1801703 https://www.tandfonline.com/doi/full/10.1080/02724634.2020.1801703
 - Rodrigo V. Pêgas, Borja Holgado & Maria Eduarda C. Leal (2019) On Targaryendraco wiedenrothi gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids, Historical Biology, doi:10.1080/08912963.2019.1690482
 













